Contact Details

Dr Caroline Buttler
National Museum Cardiff

Cathays Park
CF10 3NP

Phone: +44 (0)29 2057 3359

Staff Name

Dr Caroline Buttler

Job Title

Co-Acting Head of Natural Sciences, Head of Palaeontology

Areas of Responsibility:

Management and development of Palaeontology Section - collections, research and public engagement. Management of Mollusca Section.

Qualifications, memberships and relevant positions

BSc (Hons) Geology (University of Liverpool), MSc Palaeontology & Ecology (University of Manitoba), PhD Palaeontology (University of Wales, Swansea)

Accredited Conservator (Institute of Conservation).

Vice-President (Palaeontographical Society), Editor (Proceedings of the Geologist’s Association), Committee Member (University of Birmingham Lapworth Museum Committee).

Research Interests

My research is focused on Palaeozoic bryozoans, especially the order Trepostomata. Of particular interest is the systematics and taxonomy of this group of colonial animals, as well as their biogeography and ecology. Recent projects include the investigation of Silurian bryozoan recovery faunas after the Ordovician mass extinction, a review of bryozoan and gastropod symbiosis in the fossil record and the revision of the trepostome section for the Treatise on Invertebrate Paleontology.

I am also interested in the conservation and care of geological material in museum collections.


Palaeontology, fossils, bryozoans, trepostomes, taxonomy, biogeography, ecology, Palaeozoic, conservation, collections care


Selected Publications

Ma, J-Y, Taylor, P.D. & Buttler, C.J. 2019. Unusual compound zooecia in the trepostome bryozoan Eostenopora from the Devonian of Guizhou, China. Palaeoworld, 28, 289-294.

Buttler, C. J. Wilson M. A. 2018. Paleoecology of an Upper Ordovician submarine cave-dwelling bryozoan fauna and its exposed equivalents in northern Kentucky, USA. Journal of Paleontology, 92, 568 – 576.

Wyse Jackson, P. N, & Buttler, C. J. 2015. Preparation, Imaging, and Conservation of Paleozoic Bryozoans for Study. Invertebrate Treatise on Paleontology Online 63: Part G, Revised, Vol. 2, Chapter 3 

Popov, L., Hairapetian, V., Ghobadi Pour, M., Buttler, C., Evans, D. H., Hejazi, S. H., Jahangir, H. 2014 Llandovery fauna of Iran during the post-extinction recovery. The Third International Symposium of the International Geosciences Programme Project 589 (IGCP- 589), 105-110. 

Ma J, Buttler, C.J. & Taylor, P. D. 2014. Cladistic analysis of the ‘trepostome’ Suborder Esthonioporina and the systematics of Palaeozoic bryozoans. Studi Trentini di Scienze Naturali. 94, 153-161

Buttler, C.J., Wyse Jackson, P.N., Ernst, A. & McKinney, F.K. 2013. A Review of the early Palaeozoic biogeography of bryozoans. In Harper, D. A.T. & Servais, T. (eds) Early Palaeozoic Palaeobiogeography and palaeogeography, Geological Society, London, Memoirs, 38, 145–155.

Ernst, A & Buttler, C. 2012. Cystoporate bryozoans from the Lower – Middle Devonian of NW Spain. Neues Jahrbuch für Geologie und Paläontologie, 263, 261-285

Buttler, C. J., Wyse Jackson, P. N. & Key, M. M. Jr. 2008. Bryozoa from the Ordovician (Caradoc) of Courtown, Co. Wexford, Ireland, 9-18. In: S. J. Hageman, M. M. Key, Jr, & J. E. Winston (eds). Proceedings of the 14th International Bryozoology Association Conference. Special Publication Series of the Virginia Museum of Natural History Press; Martinsville.

Brenchley, P.J., Marshall, J.D., Harper, D.A.T., Buttler, C.J. & Underwood, C.J. 2006. A late Ordovician (Hirnantian) karstic surface in a submarine channel, recording glacio-eustatic sea-level changes: Meifod, Central Wales. Geological Journal, 40, 1-22.

Buttler, C. & Davis, M. (eds) 2006.

Things fall apart… Museum conservation in practice
. (National Museum Wales), 198pp.

Buttler, C. & Davis, M. 2006. The conservator in context: crossing curatorial boundaries, 30-33. In Saunders, D., Townsend, J.H. & Woodcock, S. (eds) The object in context: crossing conservation boundaries. Contributions to the Munich IIC Congress 2006.

Boardman, R.S & Buttler, C.J. 2005. Zooids and extrazooidal skeleton in the order Trepostomata (Bryozoa). Journal of Paleontology, 79, 1088–1104.

Blog Entries

by Caroline Buttler
26 January 2018